Complementary Habitat Use of the Eastern Box Turtle (Terrapene carolina) in the Georgia Piedmont. Bentley, K. Ph.D. Thesis, University of Georgia, Athens, Georgia, USA, 2015.
abstract   bibtex   
Complementary habitats are typically recognized for species that require distinctly different habitat types, such as aquatic and terrestrial. However, these complementary habitats can be more subtle for species that occupy only a single habitat “type”, such as forests. Most turtles exhibit complementary habitats, occupying aquatic habitats with females making seasonal movements upland to nest. Eastern box turtles (Terrapene carolina) are closely related to semi-aquatic Emydid aquatic turtles known to make extensive overland movements including upland migrations to complementary nesting habitats; and Eastern box turtles are known to utilize both upland and bottomland forests including river floodplain and wetland habitats. Whether Eastern box turtles utilize complementary habitats has not been examined. We used radiotelemetry to track 8 Eastern box turtles daily for 5 months in the Georgia Piedmont to find evidence of complementary habitat use. Our data show that all turtles had some if not most of their home range in bottomland, river floodplain forest; however, females also made repeated migrations between bottomland and upland forests, and a large portion of their home range including upland forest habitat. Females also had larger home ranges. Males seldom occurred outside bottomland forests and their home ranges were largely confined to the contour of the forested flood plain. Though males and females differed in both size and habitat within their home ranges, males and females had similar mean daily movement and cumulative distances moved over the 5 month period. Our results suggest that female Eastern box turtles exhibit complementary forest habitat requirements analogous to aquatic species within the same family. Based on the large home ranges encompassing areas in two habitats, and the high rates of movement, conservation considerations should focus on the vulnerability of females to repeated road crossings and interactions with suburban landscape features. In addition, both males and females require bottomland ecosystems, which are prone to degradation by pollution and invasive species. Conservation of bottomland forests may be more important for box turtles than is currently recognized.
@phdthesis{bentley_complementary_2015,
	address = {Athens, Georgia, USA},
	type = {Senior {Thesis}},
	title = {Complementary {Habitat} {Use} of the {Eastern} {Box} {Turtle} ({Terrapene} carolina) in the {Georgia} {Piedmont}},
	abstract = {Complementary habitats are	typically recognized for	species that require	distinctly different habitat types, such	as aquatic and terrestrial. However, these	complementary	habitats can be	more	 subtle for species that occupy only a single habitat “type”, such as forests. Most	turtles exhibit complementary habitats, occupying	aquatic habitats with	females making seasonal movements upland to nest. Eastern box turtles (Terrapene	carolina) are closely related to semi-aquatic Emydid aquatic	 turtles known to make extensive overland movements including upland migrations to complementary	 nesting habitats; and Eastern box turtles are known	 to utilize both upland and bottomland forests including	river	floodplain	and wetland habitats. Whether Eastern box turtles utilize complementary	 habitats has not been	examined. We used radiotelemetry	to track 8 Eastern box turtles daily for 5	 months in	the Georgia Piedmont	to find evidence of complementary habitat use. Our data	show	 that	all turtles had some if not most of their home	range in bottomland, river floodplain	forest; however, females also made repeated migrations between bottomland and upland forests, and a	large portion of	 their home range	including	upland forest habitat. Females also had larger home	ranges. Males seldom occurred	outside bottomland	forests and their	home ranges were largely confined to the	contour of the	 forested flood plain. Though males and	females differed in both	size	and habitat within	their	home ranges, males	and females had similar mean	daily movement and cumulative distances	moved over the	 5 month period. Our results suggest that female Eastern box turtles exhibit complementary forest habitat requirements analogous to aquatic species	within the same family. Based	on the large home ranges encompassing areas in two habitats, and the high rates of	movement, conservation considerations should focus on the vulnerability of females to	repeated road crossings and interactions  with suburban landscape	features. In addition, both	males and	females require	bottomland ecosystems, which	are prone to degradation by pollution and invasive species.	Conservation of bottomland forests may be more important for box turtles than is currently recognized.},
	school = {University of Georgia},
	author = {Bentley, Katherine},
	year = {2015},
}

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