Physiological intracellular crowdedness is defined by the perimeter-to-area ratio of sub-cellular compartments. Hiroi, N., Okuhara, T., Kubojima, T., Iba, K., Tabira, A., Yamashita, S., Okada, Y., Kobayashi, T. J, & Funahashi, A. Frontiers in physiology, 3:293, Frontiers, 2012.
bibtex   
@article{hiroi2012physiological,
  title={Physiological intracellular crowdedness is defined by the perimeter-to-area ratio of sub-cellular compartments},
  author={Hiroi, Noriko and Okuhara, Takahiro and Kubojima, Takeshi and Iba, Keisuke and Tabira, Akito and Yamashita, Shuji and Okada, Yasunori and Kobayashi, Tetsuya J and Funahashi, Akira},
  journal={Frontiers in physiology},
  volume={3},
  pages={293},
  year={2012},
  publisher={Frontiers}
}

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