From The Cover: Cross-modal integration in a dart-poison frog. Narins, P. M, Grabul, D. S, Soma, K. K, Gaucher, P., & Hödl, W. Proc Natl Acad Sci U S A, 102(7):2425-9, 2005.
doi  abstract   bibtex   
The mechanisms by which the brain binds together inputs from separate sensory modalities to effect a unified percept of events are poorly understood. This phenomenon was studied in males of the dart-poison frog Epipedobates femoralis. These animals physically and vigorously defend their territories against conspecific calling intruders. In prior field studies with an electromechanical model frog, we were able to experimentally evoke this aggressive behavior only when an auditory cue (advertisement call) was presented simultaneously with a visual cue (vocal-sac pulsations). In the present field experiments, we used a modified version of the electromechanical model frog to present territorial males with visual and auditory cues separated by experimentally introduced temporal delays or spatial disparities to probe temporal and spatial integration in this animal. In temporal integration experiments, bimodal stimuli with temporal overlap during calling bouts consistently evoked aggressive behavior; stimuli lacking bimodal temporal overlap were relatively ineffective at the same task. In spatial integration studies, despite presenting the components of the bimodal stimulus with an initial spatial disparity of up to 12 cm, fighting behavior persisted. These results demonstrate that temporal and spatial integration may be reliably estimated in a freely behaving animal in its natural habitat and that we can use aggressive behavior in this species as an index of cross-modal integration in the field.
@Article{Narins2005,
  author   = {Peter M Narins and Daniela S Grabul and Kiran K Soma and Philippe Gaucher and Walter H\"odl},
  journal  = {Proc Natl Acad Sci U S A},
  title    = {From {T}he {C}over: {C}ross-modal integration in a dart-poison frog.},
  year     = {2005},
  number   = {7},
  pages    = {2425-9},
  volume   = {102},
  abstract = {The mechanisms by which the brain binds together inputs from separate
	sensory modalities to effect a unified percept of events are poorly
	understood. This phenomenon was studied in males of the dart-poison
	frog Epipedobates femoralis. These animals physically and vigorously
	defend their territories against conspecific calling intruders. In
	prior field studies with an electromechanical model frog, we were
	able to experimentally evoke this aggressive behavior only when an
	auditory cue (advertisement call) was presented simultaneously with
	a visual cue (vocal-sac pulsations). In the present field experiments,
	we used a modified version of the electromechanical model frog to
	present territorial males with visual and auditory cues separated
	by experimentally introduced temporal delays or spatial disparities
	to probe temporal and spatial integration in this animal. In temporal
	integration experiments, bimodal stimuli with temporal overlap during
	calling bouts consistently evoked aggressive behavior; stimuli lacking
	bimodal temporal overlap were relatively ineffective at the same
	task. In spatial integration studies, despite presenting the components
	of the bimodal stimulus with an initial spatial disparity of up to
	12 cm, fighting behavior persisted. These results demonstrate that
	temporal and spatial integration may be reliably estimated in a freely
	behaving animal in its natural habitat and that we can use aggressive
	behavior in this species as an index of cross-modal integration in
	the field.},
  doi      = {10.1073/pnas.0406407102},
  keywords = {15677318},
}

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