Haul-Out Selection by Pacific Harbor Seals (phoca Vitulina Richardii): Isolation and Perceived Predation Risk. Nordstrom, C. A. Marine Mammal Science, 18(1):194–205, January, 2002.
Haul-Out Selection by Pacific Harbor Seals (phoca Vitulina Richardii): Isolation and Perceived Predation Risk [link]Paper  doi  abstract   bibtex   
The potential for non-aquatic predators to influence habitat use by harbor seals (Phoca vitulina) in a nearshore marine environment was studied by examining haul-out site use and through an experimental approach. Distance from shore, distance to possible foraging depths, peripheral water depth, and haul-out areas were quantified for each haul-out. There was a positive relationship between the number of seals hauled out and the distance from shore for eight known haul-out sites. The hypothesis that harbor seals increasingly hauled out farther offshore to reduce predation risk was tested experimentally by measuring their response to a model of a potential terrestrial predator in comparison to a control object, and to disturbance by a human at one of the study sites. Harbor seals abandoned the haul-out in the presence of the predator model, but showed little response to the controls, suggesting they possess a threat image for terrestrial predators and avoid hauling out when it is perceived. These results support the hypothesis that harbor seals select isolated sites to reduce exposure to terrestrial carnivores.
@article{nordstrom_haul-out_2002,
	title = {Haul-{Out} {Selection} by {Pacific} {Harbor} {Seals} (phoca {Vitulina} {Richardii}): {Isolation} and {Perceived} {Predation} {Risk}},
	volume = {18},
	issn = {1748-7692},
	shorttitle = {Haul-{Out} {Selection} by {Pacific} {Harbor} {Seals} (phoca {Vitulina} {Richardii})},
	url = {http://onlinelibrary.wiley.com/doi/10.1111/j.1748-7692.2002.tb01028.x/abstract},
	doi = {10.1111/j.1748-7692.2002.tb01028.x},
	abstract = {The potential for non-aquatic predators to influence habitat use by harbor seals (Phoca vitulina) in a nearshore marine environment was studied by examining haul-out site use and through an experimental approach. Distance from shore, distance to possible foraging depths, peripheral water depth, and haul-out areas were quantified for each haul-out. There was a positive relationship between the number of seals hauled out and the distance from shore for eight known haul-out sites. The hypothesis that harbor seals increasingly hauled out farther offshore to reduce predation risk was tested experimentally by measuring their response to a model of a potential terrestrial predator in comparison to a control object, and to disturbance by a human at one of the study sites. Harbor seals abandoned the haul-out in the presence of the predator model, but showed little response to the controls, suggesting they possess a threat image for terrestrial predators and avoid hauling out when it is perceived. These results support the hypothesis that harbor seals select isolated sites to reduce exposure to terrestrial carnivores.},
	language = {en},
	number = {1},
	urldate = {2015-08-13},
	journal = {Marine Mammal Science},
	author = {Nordstrom, Chad A.},
	month = jan,
	year = {2002},
	keywords = {Phoca vitulina richardii},
	pages = {194--205},
}

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