Humboldt's Enigma: What Causes Global Patterns of Mountain Biodiversity?. Rahbek, C., Borregaard, M. K., Colwell, R. K., Dalsgaard, B., Holt, B. G., Morueta-Holme, N., Nogues-Bravo, D., Whittaker, R. J., & Fjelds\aa, J. Science, 365(6458):1108–1113, September, 2019.
doi  abstract   bibtex   
Mountains contribute disproportionately to the terrestrial biodiversity of Earth, especially in the tropics, where they host hotspots of extraordinary and puzzling richness. With about 25% of all land area, mountain regions are home to more than 85% of the world's species of amphibians, birds, and mammals, many entirely restricted to mountains. Biodiversity varies markedly among these regions. Together with the extreme species richness of some tropical mountains, this variation has proven challenging to explain under traditional climatic hypotheses. However, the complex climatic characteristics of rugged mountain regions differ fundamentally from those of lowland regions, likely playing a key role in generating and maintaining diversity. With ongoing global changes in climate and land use, the role of mountains as refugia for biodiversity may well come under threat.
@article{rahbekHumboldtEnigmaWhat2019,
  title = {Humboldt's Enigma: What Causes Global Patterns of Mountain Biodiversity?},
  shorttitle = {Humboldt's Enigma},
  author = {Rahbek, Carsten and Borregaard, Michael K. and Colwell, Robert K. and Dalsgaard, Bo and Holt, Ben G. and {Morueta-Holme}, Naia and {Nogues-Bravo}, David and Whittaker, Robert J. and Fjelds{\aa}, Jon},
  year = {2019},
  month = sep,
  volume = {365},
  pages = {1108--1113},
  issn = {0036-8075, 1095-9203},
  doi = {10.1126/science.aax0149},
  abstract = {Mountains contribute disproportionately to the terrestrial biodiversity of Earth, especially in the tropics, where they host hotspots of extraordinary and puzzling richness. With about 25\% of all land area, mountain regions are home to more than 85\% of the world's species of amphibians, birds, and mammals, many entirely restricted to mountains. Biodiversity varies markedly among these regions. Together with the extreme species richness of some tropical mountains, this variation has proven challenging to explain under traditional climatic hypotheses. However, the complex climatic characteristics of rugged mountain regions differ fundamentally from those of lowland regions, likely playing a key role in generating and maintaining diversity. With ongoing global changes in climate and land use, the role of mountains as refugia for biodiversity may well come under threat.},
  copyright = {Copyright \textcopyright{} 2019, American Association for the Advancement of Science. http://www.sciencemag.org/about/science-licenses-journal-article-reuseThis is an article distributed under the terms of the Science Journals Default License.},
  journal = {Science},
  keywords = {~INRMM-MiD:z-CDLEGSHG,abiotic-factors,array-of-factors,biodiversity,ecological-domains,ecological-zones,global-scale,mountainous-areas,review,size-effect,species-diversity,species-richness},
  language = {en},
  lccn = {INRMM-MiD:z-CDLEGSHG},
  number = {6458}
}

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