The lateral inhibition of perceived contrast is indifferent to on-center/off-center segregation, but specific to orientation. Solomon, J., Sperling, G., & Chubb, C. vision research, 33(18):2671-2683, 1993.
bibtex   
@article{ Solomon_etal93,
  author = {Solomon, J.A. and Sperling, G. and Chubb, C.},
  title = {The lateral inhibition of perceived contrast is indifferent to on-center/off-center
	segregation, but specific to orientation},
  journal = {vision research},
  year = {1993},
  volume = {33},
  pages = {2671-2683},
  number = {18},
  en_number = {2.4:84}
}

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