Intergroup Selection in the Evolution of Social Systems. Wynne-Edwards, V. C. Nature, 200(4907):623--626, 1963.
Intergroup Selection in the Evolution of Social Systems [link]Paper  doi  bibtex   
@article{wynne-edwards_intergroup_1963,
	title = {Intergroup {Selection} in the {Evolution} of {Social} {Systems}},
	volume = {200},
	url = {http://dx.doi.org/10.1038/200623a0},
	doi = {10.1038/200623a0},
	number = {4907},
	urldate = {2008-07-15TZ},
	journal = {Nature},
	author = {Wynne-Edwards, V. C.},
	year = {1963},
	keywords = {group selection},
	pages = {623--626}
}

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