Cortical Neural Stem Cell Lineage Progression Is Regulated by Extrinsic Signaling Molecule Sonic Hedgehog. Zhang, Y., Liu, G., Guo, T., Liang, X. G, Du, H., Yang, L., Bhaduri, A., Li, X., Xu, Z., Zhang, Z., Li, Z., He, M., Tsyporin, J., Kriegstein, A. R, Rubenstein, J. L, Yang, Z., & Chen, B. Cell Rep, 30(13):4490–4504.e4, March, 2020.
abstract   bibtex   
Neural stem cells (NSCs) in the prenatal neocortex progressively generate different subtypes of glutamatergic projection neurons. Following that, NSCs have a major switch in their progenitor properties and produce $γ$-aminobutyric acid (GABAergic) interneurons for the olfactory bulb (OB), cortical oligodendrocytes, and astrocytes. Herein, we provide evidence for the molecular mechanism that underlies this switch in the state of neocortical NSCs. We show that, at around E16.5, mouse neocortical NSCs start to generate GSX2-expressing (GSX2(+)) intermediate progenitor cells (IPCs). In vivo lineage-tracing study revealed that GSX2(+) IPC population gives rise not only to OB interneurons but also to cortical oligodendrocytes and astrocytes, suggesting that they are a tri-potential population. We demonstrated that Sonic hedgehog signaling is both necessary and sufficient for the generation of GSX2(+) IPCs by reducing GLI3R protein levels. Using single-cell RNA sequencing, we identify the transcriptional profile of GSX2(+) IPCs and the process of the lineage switch of cortical NSCs.
@ARTICLE{Zhang2020-ie,
  title    = "Cortical Neural Stem Cell Lineage Progression Is Regulated by
              Extrinsic Signaling Molecule Sonic Hedgehog",
  author   = "Zhang, Yue and Liu, Guoping and Guo, Teng and Liang, Xiaoyi G and
              Du, Heng and Yang, Lin and Bhaduri, Aparna and Li, Xiaosu and Xu,
              Zhejun and Zhang, Zhuangzhi and Li, Zhenmeiyu and He, Miao and
              Tsyporin, Jeremiah and Kriegstein, Arnold R and Rubenstein, John
              L and Yang, Zhengang and Chen, Bin",
  abstract = "Neural stem cells (NSCs) in the prenatal neocortex progressively
              generate different subtypes of glutamatergic projection neurons.
              Following that, NSCs have a major switch in their progenitor
              properties and produce $\gamma$-aminobutyric acid (GABAergic)
              interneurons for the olfactory bulb (OB), cortical
              oligodendrocytes, and astrocytes. Herein, we provide evidence for
              the molecular mechanism that underlies this switch in the state
              of neocortical NSCs. We show that, at around E16.5, mouse
              neocortical NSCs start to generate GSX2-expressing (GSX2(+))
              intermediate progenitor cells (IPCs). In vivo lineage-tracing
              study revealed that GSX2(+) IPC population gives rise not only to
              OB interneurons but also to cortical oligodendrocytes and
              astrocytes, suggesting that they are a tri-potential population.
              We demonstrated that Sonic hedgehog signaling is both necessary
              and sufficient for the generation of GSX2(+) IPCs by reducing
              GLI3R protein levels. Using single-cell RNA sequencing, we
              identify the transcriptional profile of GSX2(+) IPCs and the
              process of the lineage switch of cortical NSCs.",
  journal  = "Cell Rep",
  volume   =  30,
  number   =  13,
  pages    = "4490--4504.e4",
  month    =  mar,
  year     =  2020,
  keywords = "Gli3; Gsx2; Shh; cerebral cortex; neural stem cells; olfactory
              bulb interneurons; oligodendrocytes",
  language = "en"
}

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